''Paraptenodytes brodkorbi'' is a proposed, but possibly invalid,

species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

penguin Penguins (order (biology), order List of Sphenisciformes by population, Sphenisciformes , family (biology), family Spheniscidae ) are a group of Water bird, aquatic flightless birds. They live almost exclusively in the Southern Hemisphere: on ...

in the genus ''

Paraptenodytes ''Paraptenodytes'' is an extinct genus of penguins which contains two or three species sized between a Magellanic penguin and an emperor penguin (''Aptenodytes forsteri''). They are known from fossil bones ranging from a partial skeleton and some ...

''.''Paraptenodytes brodkorbi''
at

Fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals ...

.org The bird was probably about the size of a

king penguin The king penguin (''Aptenodytes patagonicus'') is the second largest species of penguin, smaller, but somewhat similar in appearance to the emperor penguin. There are two subspecies: ''A. p. patagonicus'' and ''A. p. halli''; ''patagonicus'' i ...

. Known material is limited to a single

humerus The humerus (; ) is a long bone in the arm that runs from the shoulder to the elbow. It connects the scapula and the two bones of the lower arm, the radius and ulna, and consists of three sections. The humeral upper extremity consists of a roun ...

, Early

Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...

in age, found in the

Monte León Formation Monte may refer to: Places Argentina * Argentine Monte, an ecoregion * Monte Desert * Monte Partido, a ''partido'' in Buenos Aires Province Italy * Monte Bregagno * Monte Cassino * Montecorvino (disambiguation) * Montefalcione Portugal * M ...

near

Puerto San Julián Puerto San Julián, also known historically as Port St. Julian, is a natural harbour in Patagonia in the Santa Cruz Province of Argentina located at . In the days of sailing ships it formed a stopping point, south of Puerto Deseado (''Port Desir ...

in Santa Cruz Province,

Argentina Argentina (), officially the Argentine Republic ( es, link=no, República Argentina), is a country in the southern half of South America. Argentina covers an area of , making it the second-largest country in South America after Brazil, th ...

. It exists as an unnumbered specimen in the collection of the Museo Argentino de Ciencias Naturales.

Description

This

taxon In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...

is the product of a nomenclatorial dispute.

Florentino Ameghino Florentino Ameghino (born Giovanni Battista Fiorino Giuseppe Ameghino September 19, 1853 – August 6, 1911) was an Argentine naturalist, paleontologist, anthropologist and zoologist, whose fossil discoveries on the Argentine Pampas, especially ...

in 1905 described some penguin bones which he thought to be specifically distinct as ''Isotremornis nordenskjöldi'': a

tarsometatarsus The tarsometatarsus is a bone that is only found in the lower leg of birds and some non-avian dinosaurs. It is formed from the fusion of several bones found in other types of animals, and homologous to the mammalian tarsus (ankle bones) and meta ...

, a humerus, and a part of a

femur The femur (; ), or thigh bone, is the proximal bone of the hindlimb in tetrapod vertebrates. The head of the femur articulates with the acetabulum in the pelvic bone forming the hip joint, while the distal part of the femur articulates with ...

. Subsequently, the tarsometatarsus and the femur piece turned out to be from ''

Paraptenodytes antarcticus ''Paraptenodytes'' is an extinct genus of penguins which contains two or three species sized between a Magellanic penguin and an emperor penguin (''Aptenodytes forsteri''). They are known from fossil bones ranging from a partial skeleton and some ...

''.

George Gaylord Simpson George Gaylord Simpson (June 16, 1902 – October 6, 1984) was an American paleontologist. Simpson was perhaps the most influential paleontologist of the twentieth century, and a major participant in the Modern synthesis (20th century), modern ...

(1946) and

Pierce Brodkorb William Pierce Brodkorb (September 29, 1908, Chicago – July 18, 1992, Gainesville, Florida) was an American ornithologist and paleontologist. Interested in birds since childhood, he was taught to prepare birds at the age of 16. Later, he rec ...

(1963) argued about whether the bones could all be considered

syntypes In biological nomenclature, a syntype is any one of two or more biological types that is listed in a description of a taxon where no holotype was designated. Precise definitions of this and related terms for types have been established as part of ...

or whether only the wrongly assigned tarsometatarsus was designated as the

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several ...

. Brodkorb argued for the latter, and Simpson "reluctantly" agreed; ''Isotremornis nordenskjöldi'' became a

junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linna ...

of ''P. antarcticus''. Thus, another name had to be given - and added to the already long and confusing list of valid and invalid fossil penguin taxa - to the distinct humerus of the new species Ameghino had thought he described. Tongue-in-cheek, Simpson (1972) dedicated the new binomen to Brodkorb. Acosta Hospitaleche (2005) considered the humerus to be assignable to '' Paraptenodytes robustus''; Bertelli ''et al.'' (2006) disagree, but believe that it belongs into a different

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus com ...

Description

References

Further reading